It will allow you to add it to the queue (even if there are no other buildings in the queue already). However when the builder gets to the site, the construction will not begin, and the queue slot will have a red exclamation mark with the icon of the missing resource. If you then go and get that resource in any way (trade,production, moving from star, etc), the construction begins immediately.
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When it comes to Gene pools and shallow ends they can be found at the bar drinking pina colada's
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